Taxonomic Notes: Hordeum marinum Huds. is a tertiary wild relative of Barley, H. vulgare L. (USDA, ARS, National Genetic Resources Program ). Differs from Hordeum leporinum and Hordeum glaucum in that the junction of the with line drawing: Background and Aims Hordeum marinum. is a species complex that includes the diploid subspecies marinum and both diploid and tetraploid.

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Note that two ideograms were needed to include the diversity found within marinum accessions. Rapid cytological diploidization in newly formed allopolyploids of the wheat Aegilops – Triticum group. The probe labelling procedures and FISH conditions were the same as those described in earlier work de Bustos et al.

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A third locus with no counterpart in marinum was located in a subterminal position on the long arm of the satellitized chromosome pair Fig. Author information Article notes Copyright and License information Disclaimer. The first step in inferring relationships between species is the discrimination of homology among the chromosomes of related taxa. Fluorescence in situ hybridization polymorphism using two repetitive DNA clones in different cultivars of wheat.

Following the format used in previous descriptions, the chromosomes 1—7 were arranged in order of decreasing length with the satellitized chromosome at the end. Phylogenetic analyses of the diploid species of Hordeum Poaceae and a revised classification of the genus.

It remains unknown whether marinum was the second genome donor. As in marinumAAC 5 produced a rich hybridization pattern of in situ signals. One of these groups, the members of which carry the Xa genome, includes the sea barleys.

Dispersal of spikelets is probably assisted by flooding, as winter flood-lines around pools are often marked the following summer by narrow hanks of H. Phylogenetic analysis of the genus Hordeum using repetitive DNA sequences.

Hordeum marinum – Wikipedia

Combined ecological niche modelling and molecular phylogeography revealed the evolutionary history of Hordeum marinum Poaceae —niche differentiation, loss of genetic diversity, and speciation in Mediterranean Quaternary refugia. Chromosomes of each karyotype were chosen from the same metaphase. Wang H, Sun G. An annual of barish places by the sea, marinumm the trampled margins hodeum dried-up pools and ditches in grazing jarinum, on tracks and sea walls, and in the uppermost parts of saltmarshes; also, very locally, beside salt-treated roads inland.


Canadian Journal of Genetics and Cytology. Analysis of the karyotypes Fig. Published online Nov 5. The usefulness of SSR sequences as chromosomal markers revealed by ND-FISH in different species, including several members of the tribe Triticeaehas been reported previously by our group Cuadrado et al.

Note that the PDF version is the booklet as published, whereas the Excel spreadsheet incorporates subsequent corrections. The taxonomic treatment of marinum and hordfum as subspecies is generally accepted.

Nevertheless, the present results point to the existence of substantial cytogenetic differences within marinumsuggesting a less close genomic relationship between the populations of this subspecies than that which exists within the subspecies gussoneanum.

Molecular phylogeny and reticulate origins of several American polyploid Hordeum species. Chromosome identities marinmu indicated in white lettering. ACT 5 and ATC 5 returned very few signals, always of weak intensity, that could not be used as diagnostic landmarks data not shown.

This is supported by the observation of regular meiotic behaviour in hybrids between marinum and diploid forms of gussoneanum e. Atlas text references Atlas c Atlas of north European vascular plants north horddeum the Tropic of Cancer. Open in a separate window. AAG 5which produced the clearest pattern of signals, was chosen as the diagnostic probe for identifying chromosomes and for analysing the physical mapping of the set of probes in marinun in situ experiments Fig.

Hordeum marinum

Root tips were obtained from seedlings and exceptionally from plants grown in pots in a greenhouse. The tetraploid cytotype of gussoneanum overlaps with the diploids only in the farthest eastern Mediterranean, extending from there towards the east into Asia Jakob et al.

List of the Hordeum marinum material studied. Atlas c Atlas of north European vascular plants north marinhm the Tropic of Cancer. This agrees with the extensive intra- and interspecific polymorphism observed in other Triticeae species; this SSR is very abundant in barley and wheat but restricted to a few locations in H. The aim of the present work was to identify chromosomal markers for characterizing the karyotypes of a representative sample of H.


July Mean Temperature Celsius: This provided a marker of choice for distinguishing this chromosome Fig. Two subgenomes were detected in the tetraploids. If diploid gussoneanum and marinum were involved in the origin of tetraploid gussoneanumtheir chromosomes should be present in the latter.

In future, the identities of H. Count of 10km squares in Great Britain: It is noteworthy that no other diploid Hordeum species has a 5SrDNA locus on the long arm of horfeum satellitized chromosomes de Bustos et al. Metaphase accumulations and chromosome preparations were performed as previously described Cuadrado and Jouve, A case in point from Hordeum Poaceae Cladistics.

Another problem with the complex is the taxonomic status of the different cytological forms of gussoneanum. In a few places it also occurs on the upper marunum of salt-marshes, especially where these are grazed and trampled by cattle or sheep: However, some chromosomes could not be easily distinguished from others with similar patterns compare Fig.

For example, changes in the number of rDNA sites to more or less amrinum sum of those possessed by their ancestors played a major role in Triticeae evolution Ozkan and Feldman, ; Han et al.

The barley genome and its relationship with the wheat genomes. The 45SrDNA probe hybridized with two pairs of chromosomes, providing signals of different intensity. This confirms the putative involvement of gussoneanum as a diploid ancestor of H. At the intraspecific level, the conserved distribution patterns of the 2 x gussoneanum accessions contrast with the diversity observed for maarinum accessions of marinum.